Habitats & Bird Species
The bird habitats of the Indian subcontinent
can
be roughly divided into forest, scrub, wetlands (inland and littoral), marine,
grassland, desert, and agricultural land. There is some overlap between habitats:
for example, mangrove forest can also be considered as wetland, as can seasonally
flooded grassland. Many bird species require mixed habitat types.
Forests
There is a great variety of forest types in the region. Tropical forest ranges
from coastal mangroves to wet, dense evergreen forest, dry deciduous forest
and open-desert thorn forest. In the Himalayas, temperate forest includes habitats
of mixed broadleaves, moist oak and rhododendron, and dry coniferous forest
of pines and firs; higher up, subalpine forest of birch, rhododendron and juniper
occurs.
The forest areas of the region are vitally important for many of its birds.
Over half of the bird species in the subcontinent identified by Birdlife International
as globally threatened and two-thirds of the region's endemic birds are dependent
on forest.
Primary tropical and subtropical broad leaved evergreen forest
supports
the greatest diversity of bird species. Significant areas of these forest habitats
still remain in the eastern Himalayas and adjacent hills of northeast India,
in the Western Ghats, in the Andaman and Nicobar Islands, and in Sri lanka.
They also contain a higher number of endemic and globally threatened species
than any other habitat in the region.
Tropical deciduous forest, including moist and dry sal and teak forest, riverine
forest and dry thorn forest, once covered much of the plains and lower hills
of the subcontinent. Several widespread endemic species are chiefly confined
to these habitats, including Plum-headed Parakeet Psittacula cyanocephala.
Temperate and subalpine forest grows in the Himalayas. These forest types support
a relatively high proportion of species with restricted distributions, notably
White-throated Tit Aegithalos niveogularis.
Scrub
Scrub has developed in the region where trees are unable to grow, either because
soils are poor and thin, or because they are too wet, as at the edges of wetlands
or in seasonally inundated floodplains. Scrub also grows naturally in extreme
climatic conditions, as in semidesert or at high altitudes in the Himalayas.
In addition, there are now large areas of scrubland in the region where forest
has been overexploited for fodder and fuel collection or grazing.
Relatively few birds in the subcontinent are characteristic of scrub habitats
alone, but many are found in scrub mixed with grassland, in wetlands or at forest
edges.
Wetlands
Wetlands are abundant in the region and support a rich array of waterfowl. As
well as providing habitats for breeding resident species, they include major
staging and wintering grounds for waterfowl breeding in central and northern
Asia. The region possesses a wide range of wetland types, distributed almost
throughout, including mountain glacial lakes, freshwater and brackish marshes,
large water-storage reservoirs, village tanks, saline flats and coastal mangroves
and mudflats. A total of 33 of the subcontinent's wetland bird species is globally
threatened, including Spot-billed Pelican Pelecanus philippensis.
The subcontinent's most important wetland sites include Chilika lake, a brackish
lagoon in Orissa on the east Indian coast; wetlands in the Indus valley in Pakistan;
the Sunderbans in the Ganges-Brahmaputra delta in Bangladesh and India; the
extensive seasonally flooded man-made lagoons of Keoladeo Ghana National Park;
the vast saline flats of the Ranns of Kutch in northwest India; wetlands in
the moist tropical and subtropical forest of Assam and Arunachal Pradesh; the
marshes, jheels and terai swamps of the Gangetic plain; Point Calimere and Pulicat
lake on India's east coast; the Haor basin of Sylhet and east Mymensingh in
northeast Bangladesh; and the Brahmaputra floodplain in the Assam lowlands.
Small water-storage reservoirs or tanks are a distinctive feature in India and
provide important feeding and nesting areas for a wide range of waterbirds in
some places, for example on the Deccan plateau.
Grasslands
The most important grasslands for birds in the subcontinent are theseasonally
flooded grassland occurring across the Himalayan foothills and in the floodplains
of the Indus and Brahmaputra rivers, the arid grassland of the Thar desert,
and grasslands in peninsular India, especially those in Madhya Pradesh, Maharashtra
and Karnataka. These lowland grasslands support distinctive bird communities,
with a number of specialist endemic species. Most of the region's endemic grassland
birds are seriously at risk including lesser Florican Sypheotides indica, Indian
Bustard Ardeotis nigriceps, Bristled Grassbird Chaetornis striatus and Finn's
Weaver Ploceus megarhynchus.
Desert
The Thar desert isthe largest desert in the region, covering an area of 200,000
km2 in northwest India and Pakistan. There are other extensive arid areas in
Pakistan: the hot deserts of the Chagai, a vast plain west of the main mountain
ranges of Baluchistan, and the Thai, Cholistan and Sibi deserts in central and
eastern Pakistan. The far northern mountain regions, which the monsoon winds
do not penetrate, experience a cold-desert climate. There is only one bird species,
Stoliczka's Bushchat Saxicola macrorhyncha, which is virtually endemic to the
region.
Seas
As a result of increased watching by dedicated observers, several seabirds have
been added to the region's avifauna in recent years, notably the threatened
Barau's Petrel Pterodroma baraui, which was first described for science only
in 1963. Seabird breeding colonies in the subcontinent are concentrated chiefly
in the Maldives and Lakshadweep.
Migration
The large majority (1006) of the 1300 or so species recorded in the region are
resident,
although
the numbers of some of these are augmented by winter visitors breeding farther
north. Some residents are sedentary throughout the year, while others undertake
irregular movements, either locally or more widely within the region, depending
on water conditions or food supply. Many Himalayan residents are altitudinal
migrants, the level to which they descend in winter frequently depending on
weather conditions; for instance, the Grandala summers at up to 5500 m and winters
chiefly down to 3000 m, but it has been recorded as low as 1950 m in bad weather.
A number of other residents in the subcontinent breed in the Himalayas and winter
farther south in the region, one example being the endemic Pied Thrush Zoothera
wardii, which spends the winter in Sri Lanka.
Eighteen species are exclusively summer visitors to the region. Most of these,
such as Lesser Cuckoo Cucu/us po/iocepha/us, winter in Africa. Several species
breed chiefly to the north and west of the subcontinent and extend just into
Pakistan and northwest India, (or instance European Bee-eater Merops apiaster.
Some species move southeastwards, perhaps as far as Malaysia and Indonesia;
White-throated Needletail Hirundapus caudacutus is one example.
The subcontinent attracts 159 winter visitors, some of which are also passage
migrants. There is also a small number of species (19) which are known only
as passage migrants. The winter visitors originate mostly in northern and central
Asia.
Information on migration routes in the region is still patchy, but it is believed
that many of the subcontinent's winter visitors come through Pakistan, mainly
en route to India and Sri Lanka. Ringing recoveries have shown that many winter
visitors enter the subcontinent via the Indus plains. There is less information
about migration routes in the northeast of the region, but the Brahmaputra river
and its tributaries are thought to form a flyway for birds from northeast Asia.
Increasing evidence suggests that some birds breeding in the Palearctic, mainly
non-passerines, migrate directly across the Himalayas to winter in the subcontinent.
Other birds follow the main valleys, such as those of the Kali Gandaki, Dudh
Kosi and Arun in Nepal. Birds of prey, especially Aquila eagles, have also been
found to use the Himalayas as an east-west pathway in autumn; the wintering
area of these birds is unknown. Spotwinged Starling Sarog/ossa spiloptera also
undertakes east-wesmovements along the Himalayas, and it is possible that other
species perform similar migrations.
A number of pelagic and coastal passage migrants and wintering species travel
by oceanic or coastal routes. One identified coastal flyway lies on India's
east coast, linking Point Calimere in Tamil Nadu with Chilika and Pulicat Lakes.
Migration patterns of seabirds are particularly poorly understood, but there
is now evidence that some species occur more regularly than previously thought,
especially around the time of the southwest monsoon. A few species that breed
outside the region and winter in East Africa migrate through Pakistan and northwest
India, for example Rufous-tailed Rock Thrush Montico/a saxatilis. As they occur
mainly on autumn passage, they presumably use a different route in spring.
In addition to the subcontinnt's residents, summer and winter visitors and passage
migrants, nearly 100 species of vagrant have been recorded.